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Davies Arnold Cooper has lost its second head of corporate in just nine months.
DAC's Charles Wander is set to return to his former firm, Fladgate Fielder, as the steady flow of outgoing partners shows no sign of abating.
Wander says he intends to leave DAC "some time in the new year".
DAC's former head of corporate Michael Cover joined Mishcon de Reya in August. Cover headed the department until DAC began a restructuring programme in February, in which the firm decided to focus on core areas of banking, property and litigation.
As a result of the restructuring, lawyers in DAC's London and Manchester offices were made redundant.
The most recent departures include banking litigator Paul Fallon to Warner Cranston and former managing partner Nick Rochez, who left for LeBoeuf Lamb Greene & MacRae.
Nick Sinfield, DAC's managing partner, says: "Charles has been the group head of corporate resources since February." He declines to make further comment.
A DAC spokeswoman says: "I can confirm that Charles will be leaving DAC, he has given his notice.
"He is going to Fladgate Fielder. He did his articles there, and was there for 13 years and will be returning as a partner.
"David Smellie and the rest of the corporate team are very committed to the firm and are very much in the process of recruiting."
Smellie is understood to now be the most senior partner in the corporate department.
Andrew Kaufman, corporate partner at Fladgate Fielder, says: "Wander left here as a partner and we are delighted that he's coming home.
"We are looking to expand and recruit more corporate lawyers and that's also in our business plan."
Wander is still in talks with DAC to terminate his partnership agreement.